EnhancerAtlas 2.0: an updated resource with typical enhancer annotation in 600 tissue/cell types across nine species

EnhancerAtlas 2.0

EnhancerAtlas 2.0 integrated enhancers from 12 high-throupghput technologies including P300, Histone, POLR2A, TF-binding, DHS, FAIRE, Mnase-seq, GRO-seq, STARR-seq, CAGE, CHIA-PET, and MPRA. Users could click any enhancer to view its detailed information.



Osteoblast | Homo sapiens

Cell type: Osteoblast | Tissue: Bone.

The tissue/cell type 'Osteoblast' in Homo sapiens contains 23196 enhancers. Of these enhancers, 4613 are associated with 12526 SNPs. SNPs Enhancers

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ID Coordinate SNPs TF binding sites Other Evidence
HS154-14545chr22:17556150-1755695000GeneHancer;dbSUPER
HS154-14546chr22:17566120-1756747006dbSUPER;HEDD
HS154-14547chr22:17651920-1765397004GeneHancer
HS154-14548chr22:17671800-1767257001GeneHancer
HS154-14549chr22:17679960-1768043004--
HS154-14550chr22:18043410-1804998008GeneHancer;HEDD
HS154-14551chr22:18050050-1805304003GeneHancer
HS154-14552chr22:18056270-1805712000GeneHancer
HS154-14553chr22:18110170-1811136005--
HS154-14554chr22:18121650-1812745002GeneHancer;dbSUPER;HEDD
HS154-14555chr22:18252030-1825506008dbSUPER;HEDD
HS154-14556chr22:18262430-1826722006GeneHancer;dbSUPER;HEDD
HS154-14557chr22:18404470-1840870002GeneHancer;dbSUPER;HEDD
HS154-14558chr22:18424150-1843425026GeneHancer;dbSUPER;HEDD
HS154-14559chr22:18451140-18458500012GeneHancer;dbSUPER
HS154-14560chr22:18468730-18473890013GeneHancer;dbSUPER
HS154-14561chr22:18482610-1848505003dbSUPER
HS154-14562chr22:18497540-1850704009GeneHancer;dbSUPER
HS154-14563chr22:18512330-18518520010GeneHancer;dbSUPER
HS154-14564chr22:18527230-1852799002GeneHancer;dbSUPER;HEDD
HS154-14565chr22:18538590-1854009002GeneHancer;dbSUPER
HS154-14566chr22:18543700-1854485003GeneHancer;dbSUPER
HS154-14567chr22:18547020-18553250012GeneHancer;dbSUPER;HEDD
HS154-14568chr22:18561370-1856252000dbSUPER
HS154-14569chr22:18573810-1857561008GeneHancer;dbSUPER;HEDD
HS154-14570chr22:18593640-1859541000dbSUPER;HEDD
HS154-14571chr22:18632990-1863590002GeneHancer;HEDD
HS154-14572chr22:18639490-1864032000GeneHancer
HS154-14573chr22:18640590-1864233004GeneHancer
HS154-14574chr22:18894000-1889407000dbSUPER
HS154-14575chr22:18894240-1889535000dbSUPER
HS154-14576chr22:18916380-1891750002dbSUPER;HEDD
HS154-14577chr22:19105940-1910964004GeneHancer;dbSUPER;HEDD
HS154-14578chr22:19131260-1913201000--
HS154-14579chr22:19160350-1916308015dbSUPER;HEDD
HS154-14580chr22:19164720-1916523000dbSUPER
HS154-14581chr22:19165380-1916545000dbSUPER
HS154-14582chr22:19165560-1916564003dbSUPER
HS154-14583chr22:19271270-19279120012GeneHancer;dbSUPER;HEDD
HS154-14584chr22:19433800-1943490000--
HS154-14585chr22:19435070-1943518000--
HS154-14586chr22:19484040-1948490003GeneHancer;HEDD
HS154-14587chr22:19484970-1948555001GeneHancer
HS154-14588chr22:19563970-1956556011GeneHancer
HS154-14589chr22:19650650-1965694004GeneHancer
HS154-14590chr22:19664940-19682410626GeneHancer;HEDD
HS154-14591chr22:19712300-19739220131GeneHancer;HEDD
HS154-14592chr22:19786300-1978796003GeneHancer
HS154-14593chr22:19831400-1983271001GeneHancer;dbSUPER
HS154-14594chr22:19840260-1984196002dbSUPER
HS154-14595chr22:19842060-1984218000dbSUPER
HS154-14596chr22:19864760-1986561001GeneHancer;dbSUPER
HS154-14597chr22:19865710-1986588001GeneHancer;dbSUPER
HS154-14598chr22:19865960-1986770006GeneHancer;dbSUPER;HEDD
HS154-14599chr22:19868240-1987084002GeneHancer;dbSUPER
HS154-14600chr22:19870980-19881780214GeneHancer;dbSUPER;HEDD
HS154-14601chr22:19973470-1997533005dbSUPER;HEDD
HS154-14602chr22:19978340-19989090410GeneHancer;dbSUPER;HEDD
HS154-14603chr22:20009310-2001346026GeneHancer;dbSUPER;HEDD
HS154-14604chr22:20067910-2006827003dbSUPER
HS154-14605chr22:20077780-2007839000--
HS154-14606chr22:20119580-2012091002--
HS154-14607chr22:20212220-20228900126GeneHancer;dbSUPER;HEDD
HS154-14608chr22:20230630-2023172000GeneHancer;dbSUPER
HS154-14609chr22:20231930-20236650023GeneHancer;dbSUPER;HEDD
HS154-14610chr22:20306470-2030716000--
HS154-14611chr22:20307320-2030740000--
HS154-14612chr22:20748950-2074912001--
HS154-14613chr22:20770560-2077887029GeneHancer;HEDD
HS154-14614chr22:20786280-20791860113GeneHancer;HEDD
HS154-14615chr22:20848290-2085004000dbSUPER;HEDD
HS154-14616chr22:20884140-2088886004GeneHancer;dbSUPER;HEDD
HS154-14617chr22:20902830-2090572008GeneHancer;dbSUPER;HEDD
HS154-14618chr22:20905780-2090743003GeneHancer;dbSUPER
HS154-14619chr22:21057000-2105729000--
HS154-14620chr22:21161750-2116240001GeneHancer
HS154-14621chr22:21239450-2124062000GeneHancer;dbSUPER;HEDD
HS154-14622chr22:21258260-2126086004GeneHancer;HEDD
HS154-14623chr22:21311780-2131281001dbSUPER
HS154-14624chr22:21336860-2133731000--
HS154-14625chr22:21337380-2134012002HEDD
HS154-14626chr22:21391850-2139338001GeneHancer
HS154-14627chr22:21457520-2146514008GeneHancer
HS154-14628chr22:21922670-219471401319GeneHancer;dbSUPER;HEDD
HS154-14629chr22:21996810-2199715010dbSUPER
HS154-14630chr22:21997350-2199818001GeneHancer;dbSUPER;HEDD
HS154-14631chr22:22007680-22015270118GeneHancer;dbSUPER;HEDD
HS154-14632chr22:22020420-22024200012GeneHancer;HEDD
HS154-14633chr22:22058170-22064910110GeneHancer;HEDD
HS154-14634chr22:22065200-22089900037GeneHancer;HEDD
HS154-14635chr22:22095070-22113940125GeneHancer;HEDD
HS154-14636chr22:22127270-22142540121GeneHancer;dbSUPER;HEDD
HS154-14637chr22:22162140-22189340031GeneHancer;dbSUPER;HEDD
HS154-14638chr22:22218930-2222161002GeneHancer;dbSUPER;HEDD
HS154-14639chr22:22288600-2229390007GeneHancer;dbSUPER;HEDD
HS154-14640chr22:22294050-22298260015dbSUPER;HEDD
HS154-14641chr22:22300480-2230716006dbSUPER;HEDD
HS154-14642chr22:22333710-2233681001HEDD
HS154-14643chr22:22761470-22789200030GeneHancer;dbSUPER;HEDD
HS154-14644chr22:22862190-2286297001--
HS154-14645chr22:22921440-2292216001--
HS154-14646chr22:23529610-2353107002dbSUPER;HEDD
HS154-14647chr22:23586120-2358877004GeneHancer;dbSUPER;HEDD
HS154-14648chr22:23742770-2374402001--
HS154-14649chr22:23862880-2386454009GeneHancer;dbSUPER;HEDD
HS154-14650chr22:23864760-2386559000GeneHancer;dbSUPER
HS154-14651chr22:23871760-23891660147GeneHancer;dbSUPER;HEDD
HS154-14652chr22:24058780-2405921006--
HS154-14653chr22:24088180-2409289005HEDD
HS154-14654chr22:24108850-2410956011--
HS154-14655chr22:24109780-2411002000--
HS154-14656chr22:24115170-2411602000dbSUPER
HS154-14657chr22:24123600-2412415000GeneHancer;dbSUPER
HS154-14658chr22:24129450-2413079003dbSUPER
HS154-14659chr22:24170790-24175750112GeneHancer;dbSUPER;HEDD
HS154-14660chr22:24180960-2418108004dbSUPER
HS154-14661chr22:24186200-24193880018GeneHancer;dbSUPER;HEDD
HS154-14662chr22:24200220-2420090002HEDD
HS154-14663chr22:24230080-2423156003GeneHancer;dbSUPER
HS154-14664chr22:24246640-24256760218GeneHancer;dbSUPER;HEDD
HS154-14665chr22:24408180-2440853000dbSUPER
HS154-14666chr22:24456600-2445840001GeneHancer;HEDD
HS154-14667chr22:24503220-2450434000GeneHancer;HEDD
HS154-14668chr22:24541320-24544130033GeneHancer;dbSUPER;HEDD
HS154-14669chr22:24548690-2455628007GeneHancer;dbSUPER;HEDD
HS154-14670chr22:24585080-2458681004GeneHancer;dbSUPER;HEDD
HS154-14671chr22:24593700-2459462000GeneHancer;dbSUPER
HS154-14672chr22:24666950-2467266003GeneHancer
HS154-14673chr22:24709440-2471725006HEDD
HS154-14674chr22:24765290-24807550060GeneHancer;dbSUPER;HEDD
HS154-14675chr22:24828400-2482858000dbSUPER
HS154-14676chr22:24828700-2482909000dbSUPER
HS154-14677chr22:24829700-2483424004dbSUPER;HEDD
HS154-14678chr22:24856740-2486773002GeneHancer;dbSUPER;HEDD
HS154-14679chr22:24867860-2487095001GeneHancer;dbSUPER
HS154-14680chr22:24911340-2491635002GeneHancer
HS154-14681chr22:24940050-2494288004GeneHancer;dbSUPER;HEDD
HS154-14682chr22:24979780-2498159006--
HS154-14683chr22:24985920-2498882018dbSUPER
HS154-14684chr22:25075110-25088540018GeneHancer;HEDD
HS154-14685chr22:25202310-2520240000--
HS154-14686chr22:25289580-2529118003GeneHancer;dbSUPER
HS154-14687chr22:25346040-2534741001GeneHancer;dbSUPER
HS154-14688chr22:25347980-2535053009dbSUPER;HEDD
HS154-14689chr22:25363100-25375690034GeneHancer;dbSUPER
HS154-14690chr22:25380890-2538254006GeneHancer;dbSUPER
HS154-14691chr22:25387140-2538852005GeneHancer;dbSUPER
HS154-14692chr22:25391600-2539321002GeneHancer;dbSUPER
HS154-14693chr22:25465180-2546757005GeneHancer;dbSUPER
HS154-14694chr22:25506090-2550692009dbSUPER
HS154-14695chr22:25506960-2550815002dbSUPER
HS154-14696chr22:25549980-2555110000GeneHancer;dbSUPER
HS154-14697chr22:25631260-2563272003GeneHancer
HS154-14698chr22:25798750-2580053001dbSUPER
HS154-14699chr22:25800560-2580221001dbSUPER
HS154-14700chr22:25808980-25824740029GeneHancer;dbSUPER;HEDD
HS154-14701chr22:25844250-2584560002--
HS154-14702chr22:25961120-2596301001dbSUPER
HS154-14703chr22:25978710-2597981000GeneHancer;dbSUPER
HS154-14704chr22:26141210-26144980010dbSUPER;HEDD
HS154-14705chr22:26150260-2615181006dbSUPER
HS154-14706chr22:26168480-26173540013GeneHancer;dbSUPER
HS154-14707chr22:26709870-26736400054GeneHancer;HEDD
HS154-14708chr22:26751320-26761330019GeneHancer
HS154-14709chr22:26825450-2682736003GeneHancer;HEDD
HS154-14710chr22:26908140-2690835000--
HS154-14711chr22:26945840-2695068009GeneHancer;dbSUPER
HS154-14712chr22:26983990-2698601002dbSUPER;HEDD
HS154-14713chr22:27003990-2700803003GeneHancer;dbSUPER
HS154-14714chr22:27031140-27048440260GeneHancer;dbSUPER;HEDD
HS154-14715chr22:27053620-2705927004GeneHancer;dbSUPER
HS154-14716chr22:27059360-2706142001GeneHancer;dbSUPER
HS154-14717chr22:27072440-27079300045GeneHancer;dbSUPER;HEDD
HS154-14718chr22:27113420-2711921004GeneHancer
HS154-14719chr22:27152580-2715407004GeneHancer
HS154-14720chr22:27158860-2716039006GeneHancer;HEDD
HS154-14721chr22:27191670-2719286000GeneHancer
HS154-14722chr22:27196140-27210730034GeneHancer
HS154-14723chr22:27274730-27294250026GeneHancer
HS154-14724chr22:27492050-27512660052GeneHancer;dbSUPER
HS154-14725chr22:27526770-27540870057GeneHancer;dbSUPER
HS154-14726chr22:27561680-27574200059GeneHancer
HS154-14727chr22:27580130-278935806582GeneHancer;HEDD
HS154-14728chr22:27907740-2790906000GeneHancer
HS154-14729chr22:28081810-2808734007GeneHancer;dbSUPER
HS154-14730chr22:28113280-28124060028GeneHancer;dbSUPER;HEDD
HS154-14731chr22:28383100-2838424000GeneHancer
HS154-14732chr22:28386800-2838749001GeneHancer
HS154-14733chr22:28401730-2840295000--
HS154-14734chr22:28506760-28518470020GeneHancer;HEDD
HS154-14735chr22:28828540-28849560112HEDD
HS154-14736chr22:29074730-2907560011--
HS154-14737chr22:29106050-2910789002GeneHancer;HEDD
HS154-14738chr22:29108010-2911136005GeneHancer;HEDD
HS154-14739chr22:29145640-2914722011GeneHancer;dbSUPER
HS154-14740chr22:29169210-2916971000dbSUPER
HS154-14741chr22:29169770-2917019000dbSUPER;HEDD
HS154-14742chr22:29195140-2919628002dbSUPER
HS154-14743chr22:29280060-2928221001GeneHancer
HS154-14744chr22:29426070-2942707002dbSUPER
HS154-14745chr22:29427140-2942737002dbSUPER
HS154-14746chr22:29436450-2943822002GeneHancer;dbSUPER
HS154-14747chr22:29439420-2944076000GeneHancer
HS154-14748chr22:29440880-2944148001GeneHancer
HS154-14749chr22:29469220-29480650327GeneHancer;dbSUPER;HEDD
HS154-14750chr22:29524930-2952614000GeneHancer;HEDD
HS154-14751chr22:29703180-2970404002HEDD
HS154-14752chr22:29704730-2970642004HEDD
HS154-14753chr22:29706540-2970662000--
HS154-14754chr22:29706720-2970681000--
HS154-14755chr22:29706990-2970745001--
HS154-14756chr22:29783500-2978441005dbSUPER
HS154-14757chr22:29949160-2994943002--
HS154-14758chr22:29975810-2997697000--
HS154-14759chr22:29983650-2998523001GeneHancer;HEDD
HS154-14760chr22:30000100-3000206004HEDD
HS154-14761chr22:30094590-30111340139GeneHancer;dbSUPER;HEDD
HS154-14762chr22:30116520-30123650010GeneHancer;dbSUPER;HEDD
HS154-14763chr22:30140580-3014169000GeneHancer;dbSUPER;HEDD
HS154-14764chr22:30169100-3016978000GeneHancer;dbSUPER;HEDD
HS154-14765chr22:30176220-3017880003GeneHancer;dbSUPER;HEDD
HS154-14766chr22:30194460-3019632001GeneHancer;dbSUPER;HEDD
HS154-14767chr22:30233380-3023416000dbSUPER
HS154-14768chr22:30279350-3028425004GeneHancer;dbSUPER;HEDD
HS154-14769chr22:30439720-3044114001GeneHancer;HEDD
HS154-14770chr22:30554760-3055671001GeneHancer;HEDD
HS154-14771chr22:30583800-30595830219GeneHancer;dbSUPER;HEDD
HS154-14772chr22:30599640-30609780214GeneHancer;dbSUPER;HEDD
HS154-14773chr22:30613420-3061475002GeneHancer;dbSUPER
HS154-14774chr22:30628120-3063643007GeneHancer;dbSUPER;HEDD
HS154-14775chr22:30640050-3064074000dbSUPER;HEDD
HS154-14776chr22:30640920-30642660018dbSUPER
HS154-14777chr22:30647840-3065454009dbSUPER;HEDD
HS154-14778chr22:30669200-30676940310GeneHancer;dbSUPER;HEDD
HS154-14779chr22:30683550-3068469000dbSUPER
HS154-14780chr22:30684770-3068537001dbSUPER
HS154-14781chr22:30719900-3072266008dbSUPER;HEDD
HS154-14782chr22:30782750-3078301000--
HS154-14783chr22:30793160-3079518001dbSUPER
HS154-14784chr22:30813650-3081661006dbSUPER;HEDD
HS154-14785chr22:30821940-3082270001dbSUPER
HS154-14786chr22:30822830-3082315001dbSUPER
HS154-14787chr22:30840590-3084151001GeneHancer;dbSUPER
HS154-14788chr22:30936930-3094079003GeneHancer;HEDD
HS154-14789chr22:30968870-3097250004dbSUPER;HEDD
HS154-14790chr22:30986120-3098779002HEDD
HS154-14791chr22:31044910-3104609000GeneHancer;HEDD
HS154-14792chr22:31062650-3106344000--
HS154-14793chr22:31091540-3109239000--
HS154-14794chr22:31101580-3110312003GeneHancer;HEDD
HS154-14795chr22:31218700-3122954009GeneHancer;dbSUPER;HEDD
HS154-14796chr22:31264480-3126557002GeneHancer;HEDD
HS154-14797chr22:31432320-3143377001GeneHancer
HS154-14798chr22:31434550-3143621002GeneHancer;HEDD
HS154-14799chr22:31448800-31449430017GeneHancer
HS154-14800chr22:31450910-3145229003GeneHancer;HEDD
HS154-14801chr22:31471810-3147228000GeneHancer;dbSUPER
HS154-14802chr22:31481280-3148392003dbSUPER;HEDD
HS154-14803chr22:31501950-3150336003dbSUPER
HS154-14804chr22:31519110-3152012001--
HS154-14805chr22:31543150-3154851006GeneHancer;HEDD
HS154-14806chr22:31568580-3157075003GeneHancer;HEDD
HS154-14807chr22:31608410-3160945000dbSUPER
HS154-14808chr22:31644760-3165071023dbSUPER;HEDD
HS154-14809chr22:31686380-3168695000dbSUPER
HS154-14810chr22:31687070-3168716000dbSUPER
HS154-14811chr22:31687280-3168826004dbSUPER
HS154-14812chr22:31709320-3170991002GeneHancer;dbSUPER
HS154-14813chr22:31795710-3179660000--
HS154-14814chr22:31884730-3188537000--
HS154-14815chr22:31892620-3189348000--
HS154-14816chr22:31926360-3192704003GeneHancer
HS154-14817chr22:31998280-3199859000GeneHancer;dbSUPER
HS154-14818chr22:31998770-3199920000GeneHancer;dbSUPER
HS154-14819chr22:32024200-3202678003dbSUPER;HEDD
HS154-14820chr22:32033370-3203492002dbSUPER;HEDD
HS154-14821chr22:32056140-32058930010dbSUPER
HS154-14822chr22:32072260-3207340004GeneHancer;dbSUPER;HEDD
HS154-14823chr22:32075850-3207733003GeneHancer
HS154-14824chr22:32151120-3215136001--
HS154-14825chr22:32266730-3226924003GeneHancer;dbSUPER;HEDD
HS154-14826chr22:32303020-32329500031GeneHancer;dbSUPER;HEDD
HS154-14827chr22:32340810-3234121001dbSUPER
HS154-14828chr22:32346340-3235212016GeneHancer;dbSUPER;HEDD
HS154-14829chr22:32353590-324340101133GeneHancer;HEDD
HS154-14830chr22:32806190-3280805001HEDD
HS154-14831chr22:32871390-3287496024dbSUPER;HEDD
HS154-14832chr22:32923990-3292486000GeneHancer;dbSUPER
HS154-14833chr22:32925000-3292977004GeneHancer;dbSUPER;HEDD
HS154-14834chr22:32929880-3293402006GeneHancer;dbSUPER;HEDD
HS154-14835chr22:32957190-329752401115GeneHancer;dbSUPER
HS154-14836chr22:33013870-33023910015GeneHancer;dbSUPER;HEDD
HS154-14837chr22:33040210-3304116003GeneHancer;dbSUPER;HEDD
HS154-14838chr22:33118570-3312054003GeneHancer;dbSUPER
HS154-14839chr22:33156460-3315884007GeneHancer
HS154-14840chr22:33184530-3318618001GeneHancer;dbSUPER;HEDD
HS154-14841chr22:33198110-33203350013GeneHancer;dbSUPER
HS154-14842chr22:33381890-3338253002GeneHancer
HS154-14843chr22:33796980-3379781000--
HS154-14844chr22:34249060-3425104004GeneHancer;dbSUPER
HS154-14845chr22:34310760-3431598003GeneHancer;dbSUPER
HS154-14846chr22:35653550-3565490002--
HS154-14847chr22:35695970-35700480212GeneHancer;dbSUPER;HEDD
HS154-14848chr22:35705640-35712340116GeneHancer;dbSUPER;HEDD
HS154-14849chr22:35726470-3572663002GeneHancer;dbSUPER
HS154-14850chr22:35745490-3574798005GeneHancer;dbSUPER;HEDD
HS154-14851chr22:35748070-3574918002GeneHancer;dbSUPER
HS154-14852chr22:35767520-35768590011dbSUPER;HEDD
HS154-14853chr22:35770480-3577205000dbSUPER;HEDD
HS154-14854chr22:35777190-3577839010--
HS154-14855chr22:35796200-3579642000dbSUPER
HS154-14856chr22:35796600-3579734000dbSUPER
HS154-14857chr22:35826800-3582810001GeneHancer
HS154-14858chr22:35839350-35852340223GeneHancer;dbSUPER;HEDD
HS154-14859chr22:35863610-3586460000GeneHancer
HS154-14860chr22:35937550-3594284007HEDD
HS154-14861chr22:35958800-35974220038HEDD
HS154-14862chr22:35984760-3598575000GeneHancer;dbSUPER
HS154-14863chr22:36033310-3603942002GeneHancer;dbSUPER;HEDD
HS154-14864chr22:36044670-3604864004dbSUPER
HS154-14865chr22:36422410-3642428003HEDD
HS154-14866chr22:36448350-3644909003GeneHancer
HS154-14867chr22:36451150-3645253000GeneHancer
HS154-14868chr22:36461200-3646285006--
HS154-14869chr22:36476650-3647754003GeneHancer;HEDD
HS154-14870chr22:36518440-365342200161GeneHancer;dbSUPER;HEDD
HS154-14871chr22:36541030-3654152005GeneHancer;dbSUPER
HS154-14872chr22:36551510-3655390005dbSUPER
HS154-14873chr22:36556120-3655671001dbSUPER
HS154-14874chr22:36591480-3659537007GeneHancer;HEDD
HS154-14875chr22:36605880-36622250124GeneHancer;HEDD
HS154-14876chr22:36633140-3663548005dbSUPER
HS154-14877chr22:36718570-36722610015GeneHancer;dbSUPER;HEDD
HS154-14878chr22:36723530-3673413012GeneHancer;dbSUPER;HEDD
HS154-14879chr22:36745300-3675009007dbSUPER;HEDD
HS154-14880chr22:36750620-36770310137dbSUPER;HEDD
HS154-14881chr22:36775450-3677664001dbSUPER;HEDD
HS154-14882chr22:36780730-3678385002dbSUPER;HEDD
HS154-14883chr22:36792780-3679573018GeneHancer;dbSUPER;HEDD
HS154-14884chr22:36815260-3681592000GeneHancer;dbSUPER
HS154-14885chr22:36840480-3684203006GeneHancer;dbSUPER;HEDD
HS154-14886chr22:36847420-3684814000dbSUPER;HEDD
HS154-14887chr22:36850690-3685205006dbSUPER
HS154-14888chr22:36853610-3685497002GeneHancer;dbSUPER;HEDD
HS154-14889chr22:36860830-3686309001GeneHancer;dbSUPER;HEDD
HS154-14890chr22:36863960-3686482000GeneHancer;dbSUPER
HS154-14891chr22:36866570-3686914004GeneHancer;dbSUPER;HEDD
HS154-14892chr22:36876880-3687761002dbSUPER
HS154-14893chr22:36902470-3690295005--
HS154-14894chr22:36923750-3692512000--
HS154-14895chr22:36930280-36956910040GeneHancer;HEDD
HS154-14896chr22:36962540-36983510129GeneHancer;HEDD
HS154-14897chr22:36983600-370984100252GeneHancer;HEDD
HS154-14898chr22:37168770-3717171002HEDD
HS154-14899chr22:37177180-37196740027GeneHancer;HEDD
HS154-14900chr22:37268470-3727169015GeneHancer;dbSUPER;HEDD
HS154-14901chr22:37274060-37283670015GeneHancer;dbSUPER;HEDD
HS154-14902chr22:37322220-3732544004GeneHancer;HEDD
HS154-14903chr22:37364210-37387150045GeneHancer;HEDD
HS154-14904chr22:37432480-37442770115GeneHancer;dbSUPER;HEDD
HS154-14905chr22:37447990-3744913001--
HS154-14906chr22:37510600-37521870121GeneHancer;dbSUPER;HEDD
HS154-14907chr22:37552830-37573550250GeneHancer;dbSUPER;HEDD
HS154-14908chr22:37582150-3758420000dbSUPER;HEDD
HS154-14909chr22:37592020-3759626002dbSUPER;HEDD
HS154-14910chr22:37637700-3764015001dbSUPER;HEDD
HS154-14911chr22:37646400-37661940161GeneHancer;dbSUPER;HEDD
HS154-14912chr22:37662780-3766366000GeneHancer;dbSUPER;HEDD
HS154-14913chr22:37668950-3767017003GeneHancer;dbSUPER;HEDD
HS154-14914chr22:37699440-3770525004GeneHancer;dbSUPER;HEDD
HS154-14915chr22:37705360-3770702003GeneHancer;dbSUPER;HEDD
HS154-14916chr22:37711390-37736680167GeneHancer;dbSUPER;HEDD
HS154-14917chr22:37738740-37748230129GeneHancer;dbSUPER;HEDD
HS154-14918chr22:37772040-37813800068GeneHancer;dbSUPER;HEDD
HS154-14919chr22:37813960-37823330025GeneHancer;dbSUPER;HEDD
HS154-14920chr22:37828510-37865100054GeneHancer;dbSUPER;HEDD
HS154-14921chr22:37876810-3788195005dbSUPER;HEDD
HS154-14922chr22:37888810-3789009002GeneHancer;dbSUPER;HEDD
HS154-14923chr22:37892010-3789245000GeneHancer
HS154-14924chr22:37892570-3789302000GeneHancer;dbSUPER;HEDD
HS154-14925chr22:37893190-3789860007GeneHancer;dbSUPER;HEDD
HS154-14926chr22:37906430-37911970232GeneHancer;dbSUPER;HEDD
HS154-14927chr22:37914120-3791497003dbSUPER
HS154-14928chr22:37920210-37925490113GeneHancer;dbSUPER;HEDD
HS154-14929chr22:37933650-37951470062GeneHancer;dbSUPER;HEDD
HS154-14930chr22:37956600-3796207012dbSUPER;HEDD
HS154-14931chr22:37962820-3796411001dbSUPER
HS154-14932chr22:37981020-3798897018GeneHancer;dbSUPER;HEDD
HS154-14933chr22:38005360-3800790003dbSUPER
HS154-14934chr22:38029570-3803068004dbSUPER
HS154-14935chr22:38035850-3803713005dbSUPER;HEDD
HS154-14936chr22:38065250-3806600000GeneHancer;HEDD
HS154-14937chr22:38071720-3807299008dbSUPER;HEDD
HS154-14938chr22:38073070-3807375002dbSUPER;HEDD
HS154-14939chr22:38082480-3808383002--
HS154-14940chr22:38134730-3813690001GeneHancer;dbSUPER;HEDD
HS154-14941chr22:38137040-3813724003GeneHancer;dbSUPER
HS154-14942chr22:38142430-3814777017dbSUPER;HEDD
HS154-14943chr22:38147840-3815088002dbSUPER;HEDD
HS154-14944chr22:38204170-3820595002GeneHancer;HEDD
HS154-14945chr22:38213360-3821438005GeneHancer
HS154-14946chr22:38239510-3823991003--
HS154-14947chr22:38302580-3830482016dbSUPER;HEDD
HS154-14948chr22:38323830-3832780002GeneHancer;dbSUPER;HEDD
HS154-14949chr22:38329120-3833056010GeneHancer;dbSUPER;HEDD
HS154-14950chr22:38408500-38409340025GeneHancer;dbSUPER
HS154-14951chr22:38412170-3841289002GeneHancer;dbSUPER
HS154-14952chr22:38429920-3843138003GeneHancer;dbSUPER
HS154-14953chr22:38441480-38448260013GeneHancer;dbSUPER;HEDD
HS154-14954chr22:38453600-3845375000dbSUPER
HS154-14955chr22:38453860-3845524001GeneHancer;dbSUPER
HS154-14956chr22:38484170-3848475000GeneHancer
HS154-14957chr22:38484970-3848510000GeneHancer
HS154-14958chr22:38494490-3850131009GeneHancer;dbSUPER;HEDD
HS154-14959chr22:38530480-3853099000GeneHancer
HS154-14960chr22:38531090-3853194002GeneHancer
HS154-14961chr22:38565480-38577670622GeneHancer;dbSUPER;HEDD
HS154-14962chr22:38599090-3860228064dbSUPER;HEDD
HS154-14963chr22:38613660-3861529000GeneHancer;dbSUPER;HEDD
HS154-14964chr22:38623990-3862578000GeneHancer;dbSUPER;HEDD
HS154-14965chr22:38634260-3863776006GeneHancer;dbSUPER;HEDD
HS154-14966chr22:38652560-3865441000GeneHancer;dbSUPER;HEDD
HS154-14967chr22:38656290-3865842002GeneHancer;dbSUPER;HEDD
HS154-14968chr22:38665550-3866850001dbSUPER
HS154-14969chr22:38678580-3868089008GeneHancer;dbSUPER;HEDD
HS154-14970chr22:38704580-38710080013dbSUPER;HEDD
HS154-14971chr22:38710180-3871316003dbSUPER;HEDD
HS154-14972chr22:38723280-3872450001GeneHancer;dbSUPER
HS154-14973chr22:38731750-3873261007GeneHancer;dbSUPER
HS154-14974chr22:38732710-3873285000GeneHancer;dbSUPER
HS154-14975chr22:38808460-3880990004GeneHancer;dbSUPER;HEDD
HS154-14976chr22:38856210-3885906002GeneHancer;dbSUPER;HEDD
HS154-14977chr22:38864330-3886913006--
HS154-14978chr22:38897560-3890195011dbSUPER;HEDD
HS154-14979chr22:39078100-3907832000--
HS154-14980chr22:39078450-3907888001--
HS154-14981chr22:39091370-3909580002dbSUPER;HEDD
HS154-14982chr22:39102150-3910484002dbSUPER;HEDD
HS154-14983chr22:39150130-3915064013dbSUPER;HEDD
HS154-14984chr22:39150710-3915127001dbSUPER
HS154-14985chr22:39159830-3916094000GeneHancer;dbSUPER;HEDD
HS154-14986chr22:39178070-3917866010dbSUPER
HS154-14987chr22:39178880-3917994001dbSUPER;HEDD
HS154-14988chr22:39181130-3918311003GeneHancer;dbSUPER;HEDD
HS154-14989chr22:39183800-3918529007GeneHancer;dbSUPER;HEDD
HS154-14990chr22:39188690-3919007001dbSUPER
HS154-14991chr22:39224520-39239230029GeneHancer;HEDD
HS154-14992chr22:39245020-39260240220GeneHancer;HEDD
HS154-14993chr22:39263210-3926748006GeneHancer;dbSUPER;HEDD
HS154-14994chr22:39267550-3926769000dbSUPER
HS154-14995chr22:39267760-3926790000dbSUPER
HS154-14996chr22:39273260-39297910234GeneHancer;dbSUPER;HEDD
HS154-14997chr22:39319380-39322250045GeneHancer;HEDD
HS154-14998chr22:39323440-3932402000GeneHancer
HS154-14999chr22:39378480-3937911000--
HS154-15000chr22:39399230-3940526028GeneHancer
HS154-15001chr22:39410380-3941103002--
HS154-15002chr22:39417540-3941854002dbSUPER;HEDD
HS154-15003chr22:39473380-3947394001dbSUPER
HS154-15004chr22:39537440-39545790119dbSUPER;HEDD
HS154-15005chr22:39545840-3954838027dbSUPER
HS154-15006chr22:39560840-39582160144GeneHancer;dbSUPER;HEDD
HS154-15007chr22:39601280-3960203000GeneHancer;dbSUPER
HS154-15008chr22:39645960-39654830119GeneHancer;dbSUPER;HEDD
HS154-15009chr22:39669020-39677770213GeneHancer;dbSUPER;HEDD
HS154-15010chr22:39688070-3968912000GeneHancer;dbSUPER
HS154-15011chr22:39703780-3970888004GeneHancer;dbSUPER;HEDD
HS154-15012chr22:39746120-3974864033dbSUPER;HEDD
HS154-15013chr22:39784170-3978530007GeneHancer;HEDD
HS154-15014chr22:39795840-3979674002dbSUPER
HS154-15015chr22:39800460-3980258009GeneHancer;dbSUPER
HS154-15016chr22:39833140-39834890179GeneHancer;dbSUPER;HEDD
HS154-15017chr22:39853560-39873280433GeneHancer;dbSUPER;HEDD
HS154-15018chr22:39888200-39893280013GeneHancer;dbSUPER;HEDD
HS154-15019chr22:39898440-3990020004dbSUPER
HS154-15020chr22:39926810-3992839001dbSUPER;HEDD
HS154-15021chr22:39928530-3992869001dbSUPER
HS154-15022chr22:40351920-4035605002GeneHancer;dbSUPER;HEDD
HS154-15023chr22:40356180-4036199009GeneHancer;dbSUPER;HEDD
HS154-15024chr22:40441000-40450780010GeneHancer;dbSUPER;HEDD
HS154-15025chr22:40502790-40513050019GeneHancer
HS154-15026chr22:40574140-4057677003dbSUPER;HEDD
HS154-15027chr22:40672210-4067308001dbSUPER
HS154-15028chr22:40705100-4070682001GeneHancer;dbSUPER;HEDD
HS154-15029chr22:40742720-4074455001dbSUPER;HEDD
HS154-15030chr22:40766690-4076758002dbSUPER
HS154-15031chr22:41022470-4103248012GeneHancer;dbSUPER;HEDD
HS154-15032chr22:41046780-4104974009GeneHancer;dbSUPER;HEDD
HS154-15033chr22:41064570-4106618002GeneHancer;HEDD
HS154-15034chr22:41075740-4107654007GeneHancer
HS154-15035chr22:41078820-4107954001GeneHancer;HEDD
HS154-15036chr22:41103920-41119490039GeneHancer;dbSUPER;HEDD
HS154-15037chr22:41214960-4121519000--
HS154-15038chr22:41347480-4134955004--
HS154-15039chr22:41408290-41429400348GeneHancer;dbSUPER;HEDD
HS154-15040chr22:41444580-4144587001GeneHancer
HS154-15041chr22:41489100-4149049001dbSUPER
HS154-15042chr22:41592600-4159327000GeneHancer
HS154-15043chr22:41601390-4160248000--
HS154-15044chr22:41613290-4161453005GeneHancer;dbSUPER
HS154-15045chr22:41681160-4168154000dbSUPER
HS154-15046chr22:41697780-4169955004dbSUPER;HEDD
HS154-15047chr22:41763600-4176613001dbSUPER;HEDD
HS154-15048chr22:41778210-4177907007dbSUPER
HS154-15049chr22:41797800-4179887006GeneHancer;dbSUPER;HEDD
HS154-15050chr22:41799580-4180038001GeneHancer;dbSUPER
HS154-15051chr22:41808520-41811160011GeneHancer;dbSUPER;HEDD
HS154-15052chr22:41836450-4184238008GeneHancer;dbSUPER;HEDD
HS154-15053chr22:41874260-4188014004GeneHancer;dbSUPER;HEDD
HS154-15054chr22:41919280-4191983000GeneHancer;dbSUPER
HS154-15055chr22:41919950-4192084002GeneHancer;dbSUPER
HS154-15056chr22:41920970-4192119001GeneHancer;dbSUPER
HS154-15057chr22:41939700-4193999000dbSUPER
HS154-15058chr22:41958810-4195992002dbSUPER;HEDD
HS154-15059chr22:41961440-4196216002dbSUPER;HEDD
HS154-15060chr22:41984480-4198572001HEDD
HS154-15061chr22:42060050-42069930016GeneHancer;HEDD
HS154-15062chr22:42076370-4207835002--
HS154-15063chr22:42078590-4208155004HEDD
HS154-15064chr22:42196770-4220487013GeneHancer;HEDD
HS154-15065chr22:42209830-42221700417GeneHancer;dbSUPER;HEDD
HS154-15066chr22:42229360-42262830541GeneHancer;dbSUPER;HEDD
HS154-15067chr22:42306560-4230721000GeneHancer;dbSUPER
HS154-15068chr22:42315670-4232103003dbSUPER;HEDD
HS154-15069chr22:42321560-4232210000dbSUPER
HS154-15070chr22:42322340-4232264001dbSUPER
HS154-15071chr22:42341310-4234191000GeneHancer;dbSUPER
HS154-15072chr22:42342010-4234242000dbSUPER
HS154-15073chr22:42342500-4234302000dbSUPER
HS154-15074chr22:42348430-4235367007GeneHancer
HS154-15075chr22:42354950-42367930021GeneHancer;HEDD
HS154-15076chr22:42394880-42415310145HEDD
HS154-15077chr22:42475960-4247792001--
HS154-15078chr22:42666030-4266724007--
HS154-15079chr22:42833400-42844800028GeneHancer;dbSUPER;HEDD
HS154-15080chr22:42896700-4289704000dbSUPER
HS154-15081chr22:42914160-4291571004dbSUPER
HS154-15082chr22:42950050-4295094001--
HS154-15083chr22:43057960-4305898001GeneHancer
HS154-15084chr22:43075960-43077400012GeneHancer;dbSUPER;HEDD
HS154-15085chr22:43083550-4308534001GeneHancer;dbSUPER;HEDD
HS154-15086chr22:43094100-4309605001GeneHancer;dbSUPER;HEDD
HS154-15087chr22:43110040-4311680077GeneHancer;dbSUPER;HEDD
HS154-15088chr22:43121880-4312424003GeneHancer;dbSUPER
HS154-15089chr22:43138740-4314013003GeneHancer
HS154-15090chr22:43143410-4314582000GeneHancer
HS154-15091chr22:43154780-4315570001GeneHancer;HEDD
HS154-15092chr22:43165520-4316654000GeneHancer;HEDD
HS154-15093chr22:43186670-4318742000GeneHancer;dbSUPER;HEDD
HS154-15094chr22:43251640-43253110021dbSUPER
HS154-15095chr22:43319790-43342900019GeneHancer;dbSUPER;HEDD
HS154-15096chr22:43388850-4339212002GeneHancer;dbSUPER;HEDD
HS154-15097chr22:43409170-43411020010dbSUPER;HEDD
HS154-15098chr22:43462690-4346386003GeneHancer
HS154-15099chr22:43484650-4348528001--
HS154-15100chr22:43538780-4353889000dbSUPER
HS154-15101chr22:43548230-43549340129dbSUPER;HEDD
HS154-15102chr22:43579160-4358289000GeneHancer;dbSUPER;HEDD
HS154-15103chr22:43681590-4368705007GeneHancer;dbSUPER;HEDD
HS154-15104chr22:44207280-4420804000--
HS154-15105chr22:44221990-4422382002GeneHancer;dbSUPER;HEDD
HS154-15106chr22:44292890-4430408008GeneHancer;HEDD
HS154-15107chr22:44319980-4432281003HEDD
HS154-15108chr22:44351480-4435332002HEDD
HS154-15109chr22:44433100-4443449000GeneHancer;dbSUPER
HS154-15110chr22:44484600-4448593001GeneHancer
HS154-15111chr22:44528890-4453177008GeneHancer
HS154-15112chr22:44539260-4454374007GeneHancer;dbSUPER;HEDD
HS154-15113chr22:44547430-4454806002GeneHancer
HS154-15114chr22:44597660-4459919002GeneHancer
HS154-15115chr22:44699910-4470100000GeneHancer;dbSUPER
HS154-15116chr22:44703210-4470863005GeneHancer;dbSUPER;HEDD
HS154-15117chr22:44713730-44751300283GeneHancer;HEDD
HS154-15118chr22:44755240-4475810000GeneHancer;HEDD
HS154-15119chr22:44758270-4476080002GeneHancer;HEDD
HS154-15120chr22:44843960-4484658002GeneHancer;dbSUPER;HEDD
HS154-15121chr22:44893690-4489392000--
HS154-15122chr22:44899010-44947510296GeneHancer;HEDD
HS154-15123chr22:44968330-449972301222GeneHancer;dbSUPER
HS154-15124chr22:45011580-45018160127GeneHancer;dbSUPER;HEDD
HS154-15125chr22:45034370-45044600023GeneHancer
HS154-15126chr22:45051230-45059420066GeneHancer
HS154-15127chr22:45080440-4508193004GeneHancer;dbSUPER;HEDD
HS154-15128chr22:45098490-4509996001dbSUPER
HS154-15129chr22:45122520-45127600014GeneHancer;dbSUPER;HEDD
HS154-15130chr22:45396220-45404420012--
HS154-15131chr22:45435880-455100001122GeneHancer;HEDD
HS154-15132chr22:45520130-4552196000GeneHancer;HEDD
HS154-15133chr22:45523040-4552474000HEDD
HS154-15134chr22:45595920-4559683001GeneHancer
HS154-15135chr22:45601900-45607830017GeneHancer;dbSUPER
HS154-15136chr22:45607990-4560814000dbSUPER
HS154-15137chr22:45621660-4562312010GeneHancer;dbSUPER;HEDD
HS154-15138chr22:45635570-4563652002dbSUPER
HS154-15139chr22:45641860-4564320006GeneHancer;dbSUPER
HS154-15140chr22:45651640-45675860145GeneHancer;dbSUPER;HEDD
HS154-15141chr22:45685020-4568906007GeneHancer;dbSUPER;HEDD
HS154-15142chr22:45691760-45700080013GeneHancer;dbSUPER;HEDD
HS154-15143chr22:45706060-45717220019GeneHancer;dbSUPER;HEDD
HS154-15144chr22:45831160-4583208001GeneHancer;HEDD
HS154-15145chr22:45846000-4584764000GeneHancer;dbSUPER;HEDD
HS154-15146chr22:45864390-45874760016GeneHancer;dbSUPER;HEDD
HS154-15147chr22:45898940-4590927008GeneHancer;dbSUPER;HEDD
HS154-15148chr22:45933410-4593468000GeneHancer;dbSUPER
HS154-15149chr22:45944700-4594620000GeneHancer;dbSUPER
HS154-15150chr22:45947050-4595034001GeneHancer;dbSUPER;HEDD
HS154-15151chr22:45983370-4598463001GeneHancer;dbSUPER;HEDD
HS154-15152chr22:45997020-46010690117GeneHancer;dbSUPER;HEDD
HS154-15153chr22:46015500-46051550470GeneHancer;HEDD
HS154-15154chr22:46068060-46085590111GeneHancer;dbSUPER;HEDD
HS154-15155chr22:46249960-463162406156GeneHancer;HEDD
HS154-15156chr22:46327250-46340920114GeneHancer;HEDD
HS154-15157chr22:46357800-46372560320GeneHancer;HEDD
HS154-15158chr22:46390620-4639130001GeneHancer;dbSUPER
HS154-15159chr22:46391420-4639486002GeneHancer;dbSUPER;HEDD
HS154-15160chr22:46406660-4641040004dbSUPER
HS154-15161chr22:46422520-4642415003GeneHancer;dbSUPER;HEDD
HS154-15162chr22:46430810-46433200017GeneHancer;dbSUPER;HEDD
HS154-15163chr22:46438520-4643944000dbSUPER;HEDD
HS154-15164chr22:46439580-4644048000dbSUPER
HS154-15165chr22:46455020-46476870227dbSUPER;HEDD
HS154-15166chr22:46487010-4649106002dbSUPER
HS154-15167chr22:46494440-4649907001GeneHancer;dbSUPER;HEDD
HS154-15168chr22:46499130-4650363017GeneHancer;dbSUPER
HS154-15169chr22:46509810-46524730018GeneHancer;dbSUPER;HEDD
HS154-15170chr22:46546560-4654779001dbSUPER
HS154-15171chr22:46547870-4654851001dbSUPER
HS154-15172chr22:46628990-4662987001GeneHancer;dbSUPER;HEDD
HS154-15173chr22:46644180-4664615002dbSUPER
HS154-15174chr22:46664220-4666441000--
HS154-15175chr22:46699160-4670029002GeneHancer;HEDD
HS154-15176chr22:46731740-4673330002dbSUPER
HS154-15177chr22:46786380-4678708000GeneHancer;dbSUPER
HS154-15178chr22:46787180-4678739000GeneHancer;dbSUPER
HS154-15179chr22:46910340-4691181012GeneHancer;dbSUPER;HEDD
HS154-15180chr22:46923240-4692952009GeneHancer;dbSUPER
HS154-15181chr22:46938070-4693974002GeneHancer;dbSUPER
HS154-15182chr22:46942560-4694425003GeneHancer;dbSUPER
HS154-15183chr22:46959200-4696051003GeneHancer;dbSUPER
HS154-15184chr22:46970900-4697558007dbSUPER;HEDD
HS154-15185chr22:46987800-4698944001GeneHancer;dbSUPER;HEDD
HS154-15186chr22:47133070-4713390000dbSUPER
HS154-15187chr22:47144710-47147280010GeneHancer;dbSUPER;HEDD
HS154-15188chr22:47149300-4715111001GeneHancer;dbSUPER;HEDD
HS154-15189chr22:47186350-4718841002GeneHancer;dbSUPER
HS154-15190chr22:47194390-4719500001GeneHancer;dbSUPER
HS154-15191chr22:47224500-4722743001GeneHancer;dbSUPER
HS154-15192chr22:47327040-4733302008GeneHancer
HS154-15193chr22:47549750-4755133002GeneHancer
HS154-15194chr22:47722050-47729600036GeneHancer
HS154-15195chr22:47732430-4773376000GeneHancer
HS154-15196chr22:50247640-5025510007GeneHancer;dbSUPER;HEDD
HS154-15197chr22:50326880-50332040021dbSUPER;HEDD
HS154-15198chr22:50340700-50349140118GeneHancer;dbSUPER;HEDD
HS154-15199chr22:50360120-5036186002dbSUPER;HEDD
HS154-15200chr22:50466110-5046699000GeneHancer
HS154-15201chr22:50547680-5055207003GeneHancer;dbSUPER;HEDD
HS154-15202chr22:50585580-5058804002GeneHancer;HEDD
HS154-15203chr22:50624430-5062738005dbSUPER;HEDD
HS154-15204chr22:50628210-5063003021dbSUPER;HEDD
HS154-15205chr22:50640020-5064112000dbSUPER
HS154-15206chr22:50688400-5068849001dbSUPER
HS154-15207chr22:50688590-5068885000dbSUPER
HS154-15208chr22:50688950-5068919003dbSUPER
HS154-15209chr22:50689340-5068940000dbSUPER
HS154-15210chr22:50698310-5069959006HEDD
HS154-15211chr22:50706380-5070860000GeneHancer;dbSUPER
HS154-15212chr22:50729030-50733140010GeneHancer;dbSUPER;HEDD
HS154-15213chr22:50733210-50745980122dbSUPER;HEDD
HS154-15214chr22:50764110-5076524000--
HS154-15215chr22:50781980-5078210000--
HS154-15216chr22:50911560-5091321001dbSUPER
HS154-15217chr22:50963210-5096387012dbSUPER
HS154-15218chr22:50976010-5098146017GeneHancer;dbSUPER;HEDD
HS154-15219chr22:51049980-51061180011GeneHancer;dbSUPER;HEDD
HS154-15220chr22:51065840-5106598000dbSUPER
HS154-15221chr22:51094630-5109542010GeneHancer;dbSUPER
HS154-15222chr22:51136140-5113711003GeneHancer;dbSUPER