EnhancerAtlas 2.0: an updated resource with typical enhancer annotation in 600 tissue/cell types across nine species
EnhancerAtlas 2.0
TagContent
EnhancerAtlas ID
DM013-00261 
Organism
Drosophila melanogaster 
Tissue/cell
L3_brain 
Coordinate
chr2L:1317336-1322052 
TF binding sites/motifs
Number: 165             
TFJASPAR IDCoordinateMotif SequenceStrand-Log10(p-value)
Abd-BMA0165.1chr2L:1318178-1318184CATAAA-4.01
AntpMA0166.1chr2L:1318575-1318581TAATGA+4.01
AntpMA0166.1chr2L:1318690-1318696TAATGA+4.01
AntpMA0166.1chr2L:1321249-1321255TAATGA+4.01
AntpMA0166.1chr2L:1317980-1317986CATTAA-4.01
B-H1MA0168.1chr2L:1317508-1317514AATTAA-4.01
B-H1MA0168.1chr2L:1318738-1318744AATTAA-4.01
B-H1MA0168.1chr2L:1321086-1321092AATTAA-4.01
B-H2MA0169.1chr2L:1317508-1317514AATTAA-4.01
B-H2MA0169.1chr2L:1318738-1318744AATTAA-4.01
B-H2MA0169.1chr2L:1321086-1321092AATTAA-4.01
C15MA0170.1chr2L:1317508-1317514AATTAA-4.01
C15MA0170.1chr2L:1318738-1318744AATTAA-4.01
C15MA0170.1chr2L:1321086-1321092AATTAA-4.01
CG11085MA0171.1chr2L:1317508-1317514AATTAA-4.01
CG11085MA0171.1chr2L:1318738-1318744AATTAA-4.01
CG11085MA0171.1chr2L:1321086-1321092AATTAA-4.01
CG15696-RAMA0176.1chr2L:1317508-1317514AATTAA-4.01
CG15696-RAMA0176.1chr2L:1318738-1318744AATTAA-4.01
CG15696-RAMA0176.1chr2L:1321086-1321092AATTAA-4.01
CG32532MA0179.1chr2L:1317508-1317514AATTAA-4.01
CG32532MA0179.1chr2L:1318738-1318744AATTAA-4.01
CG32532MA0179.1chr2L:1321086-1321092AATTAA-4.01
CG34031MA0444.1chr2L:1317508-1317514AATTAA-4.01
CG34031MA0444.1chr2L:1318738-1318744AATTAA-4.01
CG34031MA0444.1chr2L:1321086-1321092AATTAA-4.01
CG4328-RAMA0182.1chr2L:1318400-1318406TTATTG+4.01
CG4328-RAMA0182.1chr2L:1321906-1321912TTATTG+4.01
CG4328-RAMA0182.1chr2L:1321931-1321937TTATTG+4.01
CG4328-RAMA0182.1chr2L:1320928-1320934AATAAA-4.01
CTCFMA0531.1chr2L:1319385-1319399AAACAGGTGGAGAC+4.01
CTCFMA0531.1chr2L:1318803-1318817GCGCCATTTCTGCC-4.08
DMA0445.1chr2L:1320915-1320925CTATTGTTAA+4.18
DMA0445.1chr2L:1321199-1321209AAACAAAGAA-4.26
DMA0445.1chr2L:1319572-1319582AAACAATGGT-4.39
DfdMA0186.1chr2L:1318575-1318581TAATGA+4.01
DfdMA0186.1chr2L:1318690-1318696TAATGA+4.01
DfdMA0186.1chr2L:1321249-1321255TAATGA+4.01
DfdMA0186.1chr2L:1317980-1317986CATTAA-4.01
DllMA0187.1chr2L:1318752-1318758AATTGC+4.1
DllMA0187.1chr2L:1317507-1317513CAATTA-4.1
DllMA0187.1chr2L:1318737-1318743CAATTA-4.1
EcR|uspMA0534.1chr2L:1320841-1320855GCATCAATGAATTT+4.05
Eip74EFMA0026.1chr2L:1320692-1320698TTCCGG-4.01
Eip74EFMA0026.1chr2L:1321844-1321850TTCCGG-4.01
HHEXMA0183.1chr2L:1320173-1320180TCAATTA+4.06
HHEXMA0183.1chr2L:1320170-1320177AATTCAA-4.23
HmxMA0192.1chr2L:1317508-1317514AATTAA-4.01
HmxMA0192.1chr2L:1318738-1318744AATTAA-4.01
HmxMA0192.1chr2L:1321086-1321092AATTAA-4.01
KrMA0452.2chr2L:1318190-1318203TTACCTCTTTCTA+4.07
KrMA0452.2chr2L:1320576-1320589ACAAGGGGTTGTC-4.44
MadMA0535.1chr2L:1321772-1321786TGGCGACAGGACTC+4.26
MadMA0535.1chr2L:1318481-1318495CGACGAGAACGGGG+4.36
NK7.1MA0196.1chr2L:1317508-1317514AATTAA-4.01
NK7.1MA0196.1chr2L:1318738-1318744AATTAA-4.01
NK7.1MA0196.1chr2L:1321086-1321092AATTAA-4.01
ScrMA0203.1chr2L:1318575-1318581TAATGA+4.01
ScrMA0203.1chr2L:1318690-1318696TAATGA+4.01
ScrMA0203.1chr2L:1321249-1321255TAATGA+4.01
ScrMA0203.1chr2L:1317980-1317986CATTAA-4.01
UbxMA0094.2chr2L:1318817-1318824TTAATTA+4.23
Vsx2MA0180.1chr2L:1318818-1318826TAATTAAC-4.04
br(var.2)MA0011.1chr2L:1319696-1319703CCTATTT+4.12
br(var.2)MA0011.1chr2L:1317810-1317817TCTATTT+4.27
br(var.2)MA0011.1chr2L:1318199-1318206TCTATTT+4.27
br(var.3)MA0012.1chr2L:1318115-1318125AAACTAAAAA+4.87
br(var.3)MA0012.1chr2L:1318137-1318147TTTTAGTTTA-4.87
br(var.4)MA0013.1chr2L:1320976-1320986TTTTTTATAA-4.08
br(var.4)MA0013.1chr2L:1317768-1317778TCGTTTTCTA-4.25
brMA0010.1chr2L:1320725-1320738ATTTGATTTTTGT-4.01
brMA0010.1chr2L:1320927-1320940CAATAAAAAAAAA+4.02
brMA0010.1chr2L:1320893-1320906ATTTGTTATAGAT-4.09
brMA0010.1chr2L:1319350-1319363TAAAAATCAAGTT+4.32
brkMA0213.1chr2L:1320184-1320191GCGCCAC-4.24
bshMA0214.1chr2L:1321484-1321490CATTAA-4.1
btdMA0443.1chr2L:1321855-1321864GGGGGCGTA+5.1
btnMA0215.1chr2L:1318575-1318581TAATGA+4.01
btnMA0215.1chr2L:1318690-1318696TAATGA+4.01
btnMA0215.1chr2L:1321249-1321255TAATGA+4.01
btnMA0215.1chr2L:1317980-1317986CATTAA-4.01
cadMA0216.2chr2L:1321904-1321914TTTTATTGTC-4.54
cadMA0216.2chr2L:1320926-1320936GCAATAAAAA+4.67
cadMA0216.2chr2L:1319655-1319665GTCACAAAAA+4
cnc|maf-SMA0530.1chr2L:1320114-1320128CTGCCTCTGCAGAT+4.17
dl(var.2)MA0023.1chr2L:1319689-1319698TGGCTTTCC+4.11
dl(var.2)MA0023.1chr2L:1319230-1319239GAAAACCAC-4.35
dlMA0022.1chr2L:1321508-1321519CGAAACAACCG-4.13
dlMA0022.1chr2L:1321674-1321685GTGCTTTTCCA+4.14
dlMA0022.1chr2L:1318955-1318966AGAAAAAACCG-4.94
emsMA0219.1chr2L:1318575-1318581TAATGA+4.01
emsMA0219.1chr2L:1318690-1318696TAATGA+4.01
emsMA0219.1chr2L:1321249-1321255TAATGA+4.01
emsMA0219.1chr2L:1317980-1317986CATTAA-4.01
exdMA0222.1chr2L:1318646-1318653TTTGACA+4.24
exexMA0224.1chr2L:1319200-1319206TAATTA+4.01
exexMA0224.1chr2L:1319201-1319207AATTAC-4.01
fkhMA0446.1chr2L:1317727-1317737GTTTGCCTAA+6
ftzMA0225.1chr2L:1318575-1318581TAATGA+4.01
ftzMA0225.1chr2L:1318690-1318696TAATGA+4.01
ftzMA0225.1chr2L:1321249-1321255TAATGA+4.01
ftzMA0225.1chr2L:1317980-1317986CATTAA-4.01
gcm2MA0917.1chr2L:1317879-1317886CCCGCAC-4.49
gcm2MA0917.1chr2L:1320148-1320155CCCGCAT-4.65
gtMA0447.1chr2L:1318701-1318710TTACATAAT+4.48
gtMA0447.1chr2L:1318701-1318710TTACATAAT-4.48
hMA0449.1chr2L:1321261-1321270GCACACGCC+4.43
hMA0449.1chr2L:1321261-1321270GCACACGCC-4.43
hbMA0049.1chr2L:1318944-1318953AAAAAAAAA+4.35
hbMA0049.1chr2L:1318941-1318950CACAAAAAA+4.64
hbMA0049.1chr2L:1320928-1320937AATAAAAAA+4.88
hbMA0049.1chr2L:1318972-1318981CACAAAAAA+5.01
hkbMA0450.1chr2L:1318492-1318500GGGCGGGA+4.75
kniMA0451.1chr2L:1321390-1321401ATGCGGAGCAG+4.41
kniMA0451.1chr2L:1320886-1320897TGCTCTGATTT-4.82
kniMA0451.1chr2L:1319600-1319611TGCCCTGCTTT-4.9
lmsMA0175.1chr2L:1317508-1317514AATTAA-4.01
lmsMA0175.1chr2L:1318738-1318744AATTAA-4.01
lmsMA0175.1chr2L:1321086-1321092AATTAA-4.01
nubMA0197.2chr2L:1321422-1321433ATTTAAATTAT+4.63
oddMA0454.1chr2L:1319633-1319643ACAGTAGGAA+4.09
oddMA0454.1chr2L:1321864-1321874GCAGTAGCAC+4.38
oddMA0454.1chr2L:1318365-1318375TGCTTCTGTT-4.38
onecutMA0235.1chr2L:1317377-1317383TGATTT+4.01
onecutMA0235.1chr2L:1317575-1317581TGATTT+4.01
onecutMA0235.1chr2L:1320723-1320729TGATTT+4.01
onecutMA0235.1chr2L:1320728-1320734TGATTT+4.01
onecutMA0235.1chr2L:1319277-1319283AATCAA-4.01
onecutMA0235.1chr2L:1319354-1319360AATCAA-4.01
onecutMA0235.1chr2L:1321078-1321084AATCAA-4.01
opaMA0456.1chr2L:1319591-1319602CACCCCTTCTG+4.32
panMA0237.2chr2L:1321548-1321561CGGAATTTTTCGA+4.07
sdMA0243.1chr2L:1318905-1318916AAATTCATCGA+4.72
slboMA0244.1chr2L:1319726-1319733TTACACA+4.02
slboMA0244.1chr2L:1318678-1318685GTGTAAT-4.02
slboMA0244.1chr2L:1319320-1319327TGGCAAA+4.14
slboMA0244.1chr2L:1317439-1317446TTGCAAA+4.4
slboMA0244.1chr2L:1317504-1317511TTGCAAT-4.4
slboMA0244.1chr2L:1320781-1320788TTGCACA+4.74
slouMA0245.1chr2L:1317508-1317514AATTAA-4.01
slouMA0245.1chr2L:1318738-1318744AATTAA-4.01
slouMA0245.1chr2L:1321086-1321092AATTAA-4.01
slp1MA0458.1chr2L:1317489-1317499TGTTTTCCCT+4.3
snaMA0086.2chr2L:1319385-1319397AAACAGGTGGAG+4.81
snaMA0086.2chr2L:1320248-1320260TGCCCCTGTCGG-4
su(Hw)MA0533.1chr2L:1318562-1318582TATGCAGCATAATTAATGAG-4.13
su(Hw)MA0533.1chr2L:1318725-1318745CTTCAAGACATGCAATTAAA+4.42
su(Hw)MA0533.1chr2L:1320273-1320293ATTATTGCCTACTTTTCGGG-7.74
tinMA0247.2chr2L:1319257-1319266GCCGAGTGG+4.03
tinMA0247.2chr2L:1318711-1318720CACTCGAAA-4.79
tinMA0247.2chr2L:1319236-1319245CACTTGAAC-4.84
tllMA0459.1chr2L:1320749-1320758TTGACTTCG-4.01
tllMA0459.1chr2L:1319583-1319592AAAGTCAGC+4.04
tllMA0459.1chr2L:1318067-1318076AAAGCCAAG+4.25
tllMA0459.1chr2L:1317446-1317455TTGGCTTTT-4.29
tllMA0459.1chr2L:1318984-1318993TTGACTTTT-5.78
tupMA0248.1chr2L:1321484-1321490CATTAA-4.1
twiMA0249.1chr2L:1321469-1321480GCCATATGTTT+4.28
twiMA0249.1chr2L:1320535-1320546CACATATGCTA-4.38
unc-4MA0250.1chr2L:1317508-1317514AATTAA-4.01
unc-4MA0250.1chr2L:1318738-1318744AATTAA-4.01
unc-4MA0250.1chr2L:1321086-1321092AATTAA-4.01
uspMA0016.1chr2L:1320217-1320226GGGTCAGCC+4.58
vndMA0253.1chr2L:1319237-1319245ACTTGAAC-4.32
vndMA0253.1chr2L:1321014-1321022TTCAAGTA+4.4
Enhancer Sequence
CGTTAGTGTT GCGCATTAAA GAGAATTAAT TATTTAACAA TTGATTTAAA TTGTAATAGT 60
CTTAAAAATA TACATATTAA TTATACCATT CTGCATGAAG ACATTGCAAA TTGGCTTTTC 120
TTTTCAAATT TCCCTCAAAT CCTCGTTACT CCTTGTTTTC CCTCCTTTTT GCAATTAAAC 180
TTTTCTGGCG TTTCATTTAT TCCGCTTGGC TTTATTTGCT GAATTCCCGT TTTCGCATTT 240
GATTTATCAG CGCTTCTCAT TTCCGTTTTG GCCATTGATT ATCTCCATCA GCTTCTGCGT 300
CCTTTTGGTC CTTTGGCGTG CGTGTGTGTC CGTGAATCGC TGTATCCTTG TATCTCACAG 360
CTCGTTGTAA ACTCGTTCTG GCAGCTTTAA TGTTTGCCTA AGCGGAAATG GAAAACTTTT 420
GCACGTAGTC CTTCGTTTTC TAAGTGGCCA AGTAGAAGAA TGCACGTTTC AGCTTCTATT 480
TTGTTGCCCC AGCGATCGAG CATGAAAAGT TTTGGTTTCA GTTTGGCCGT TCACGTATTT 540
GCACCCGCAC ACATTCCACT GCCATGTAAC ACTTGCCAGG ATTTCTTTAA GGACATAAGC 600
GAGACTCCAG ATGGTCTGAG CTCGAATTTG CACGCACGTC TGCTCATTAA ATACGCATAG 660
AAACAAAACT GAAATGAATA GATGCTGGCT TGCCAGAGGC GCAGTTGCGA CTCCAAAAGG 720
GGGGTCCTTC AAAAGCCAAG AAGTGGCCAA TGGGAATACA GAACTGTAAC TGGAACATTA 780
AACTAAAAAT GATATACATC ATTTTAGTTT AGATTAGTAG AGCGATCTTA ATCTTAAATC 840
TTCATAAAAC ACGGTTACCT CTTTCTATTT CAAACTCGCA CTCCCTCCAA GGATTCGTAT 900
GCCCCTGTAA CTTCGAGGAC ACGCACACAC TTGGATGTAT GTTGTGCTAG ATAATGAATC 960
TTCCGATTCT CCGATCCCCC CGACCAACAA CCGCTTTTGT ATCTATCGGA TTGTCTGTCT 1020
GTGTGTGGCT GCTTCTGTTG AGCTGCTGTC TGCCTTATTT GCTTTTATTG TCTTCAACTT 1080
TATTCATATC CTGTTAAAAT GTATAACTTT GGGTTTTCGG GTTATAAATT AAATACGAGA 1140
CTATGCGACG AGAACGGGGC GGGATGGCAG TGGCCTGGAG GGAGCTTAGG CTCCTCAAAG 1200
CAGACGATTG TGTTGATAAA CTGCGATATG CAGCATAATT AATGAGCTTG TCATACAATG 1260
CCAATATTTT ACATTAAACA CACACACACA CACACAGCCG GAGAGACATG TTTGACATGT 1320
GCATTGTGTA CTTGAGGCAA ATGTGTAATG GTTTTAATGA GTAAATTACA TAATGCACTC 1380
GAAATCCTGC TTCAAGACAT GCAATTAAAT TGAGATAATT GCTTTCGATT CCCAAGCAGC 1440
CTCCACTTCT TATAGAGACA CTCAGTAGCG CCATTTCTGC CTTAATTAAC TCTTACCTGT 1500
GTTATTTGCG GTCCTTAATT TAGCGAAACT TGACAGGTCA TCAATAATTG AGATGCCAAC 1560
CAACCTGCGA AATTCATCGA AAATTGTCCA AAACGAGGAA AACTCCACAA AAAAAAAAGA 1620
GAAAAAACCG TTGGAGCACA AAAAACCTTT GACTTTTGGT TACTTTTCAT TGGCTCTCGG 1680
CAACAAAATG TGGAACAAAA ACCATTCAAG ATATCTATAG ATATTTTGAA CGATTTTTAA 1740
ATACTCTATG ACTAAAGTTT ATAGGTAAAA TAGGTAACAA TTTTCAAAAC AATCTTCCAA 1800
AATCCCAGGC CCCAGCCAAA TTAATACATA CTGGCTTTTT AACGCAACCC AAAGTGATTT 1860
AAAGTAATTA CCTCCAGTTA AGGGTATAAA TAATGAAAAC CACTTGAACA AGTCAAGTCA 1920
AGCCGAGTGG AAGACGTTGG AAATCAATAT ACACAAGCAC AGACACAAAA CCAGAAAAAG 1980
AAAATGGCAA AGTCGAAATG CAAGACGAAA ACGATAAAAA TCAAGTTCAT TTACCAAACC 2040
AAAGAGGGCA AACAGGTGGA GACACAGCCA GATACCACAC CTGGTTAGTT CGTCGTACTT 2100
TTTTAAGATC TGCTGGTTGC CTGGCATTAA AGTTAATAGA AAAAGTTGCT GTTGGCCAAG 2160
ATTCAATCAT GCGATATAAA CAAGCAAGCA CACACACACA CACACTGAGA CCGAAAAGTT 2220
GGCGGCCAGG TAAATAAAAC AATGGTCAAA GTCAGCACCC CTTCTGCCCT GCTTTTCCCC 2280
AAAGAAATTC CCTCACCACA GTAGGAAAAG CGATAAGAAG TCACAAAAAC TTTTCTCTGC 2340
TCCTTCATTT CCTTGGCTTT CCTATTTTTT GTCAGTCTCA AGTTGAAAAA TTACACAATG 2400
CAGGACAGTT GGGAGAGCTA AGTTTCTGGC CTTTCCCTAT TTGGCCGGCT GATATTTGCC 2460
GGTGTTGATT GAAATTTGCA TGTTGACTAT CAGCCAGGAA GAGCAACTCA AAGTAACTCA 2520
AATAACAACT TTTAAATTCA CCCTTAAACT AAGTTTGCAA ATTGAAATAC TCAGGCAAGG 2580
TGATCTTGAA AAGTGAAAGG AGGAGAAGGT AAAGTATGGA TACCTCGATA TTCATATATT 2640
TCATATGATA GTAAATGTTT CAGATCTATG AAAATATAAA TTAACTAAAG GAAAAGAAAT 2700
GGCTTTCTTT TAAATGCCCC ATGGCAACAT CTTTAATCTT TAAGTCGAAA TCATGCTCCT 2760
TCCATCGCAG TTGTTCCCCT GCCTCTGCAG ATTAAATCCG TTGAAGCATG TGCCCGCATA 2820
TTGAAATCTC ATTTAATTCA ATTACAAAGC GCCACAACTT AAAGCAGACA AAGGTCCTGA 2880
GGGGTCAGCC AGTGAACTCC AACCAGATTT TCTGCCCCTG TCGGAGTTTC ACTTACGATT 2940
ATTGCCTACT TTTCGGGGTG GCAGGACGGG AAGTTAAGGC GGTCCGAGCT GATGGCGAGT 3000
GTACAACACG GCTTATGCGC CACATGCAAC ATGTATGCCT CCATTTCCGA CTTCTCCTCA 3060
TGCGAACGGA TCTGCACCCC ACGTATATAC TTTACATGCT CGAATTGTCA TTGAAGCGTT 3120
TAGACATTTT ATTAGATTGT TCCAAAGACC ATCGCCAGGA TATCGCCCCT TGGCTCGACT 3180
GACTGCCCCG AGATTTATGC ACATATGCTA AGCACAGGCA ATCGAGGGGT TGTCCCATGG 3240
ACAAGGGGTT GTCTGCTGAC TCCTGGCCGT CTGAGTTTTC CGACTTCTGT TTGGTTTGCC 3300
TTCCTCCGCT TTTCTTTGGT TTCCTGCTTT TCGCCTGCCT TGGCATTTCC CCGCTTTTCC 3360
GGCAGCTTGT TCCCCCCAAA TTAAAATTGA TTTGATTTTT GTGTGCTCCC TCCTTGACTT 3420
CGGGATTTTT GCTAATACGC TGAAATTGCA CATGTGCTGG CCGGTTTGTT CTTTTCGGGC 3480
ATCCTTGGAA TTTTCGTTAA TAAGGGCATC AATGAATTTT AATTTTGAGT AAAAGGAAAA 3540
GTATGTGGTT TGCTCTGATT TGTTATAGAT AAAGTGAAGC TATTGTTAAT GCAATAAAAA 3600
AAAATATTGA AATAATATGA TTTATTAGAG CAGCGTGTGT TTTTTTATAA AAATGAAAGA 3660
AATCATTTTG TTATTGGATT CAAGTAATGC CAATATACGT TTGAGCCCTT TTACCCTCAA 3720
CCTCATTGCG TTGAGCTCCC AAAATCAATC AATTAAATGT AACCCTCTGA AGGCCCTGCT 3780
TAAAAATCTA GACACATCCC CCGTTCTAAC CAAATATCCA AAGAATTCGT GTAAATTCCC 3840
CCATAATATG CTGTTCCCCC CGAAAACAAA GAACGTTAAA CGTCTATGCA TGCAAAAATG 3900
CTAAAGATTT TTTTAATGAA GCCCGGCACA CGCCTGGCAC ACAAATCTCT CTGGGATTTG 3960
ACGAGCGAAA ATTAATACGA AATCGAAAAA TCATTTACCG TCGCTTATCG CAACGACGAC 4020
AGGCAGGTTG AAGGGCCAAC GCTGCGTATG CGCAATGCGG AGCAGAGTGG AGCTGGAAAG 4080
TAGCTTATTT AAATTATCAA CACGTTCGAT ACTCAGCTCC ACGGGTCGCC ATTGCCATAT 4140
GTTTGGTCCA TTAAGATAAG CGTCAATCGA AGCGAAACAA CCGGGAGCTG GGGACTCATC 4200
CGTGGGGTGG TTCGGAATTT TTCGAGATGG AGCGCAACTG TAAATCCGGC AAATGTGTAA 4260
AATATTGTAG GGCACGTTTT AAAATTACCA TCAATAATGC AGGCTAAACT CAGGCCGAAC 4320
AAGGAAAGCC TCGCCAGGGT GCTTTTCCAC CTAGATTTCC ACCGCTTTCC CAGTCCTCCA 4380
TTTCCCAGTC GATGTCAGTG ACACTTGGCC CGGCCCTTCG ACTTTTGTTT AAGTCCTGGC 4440
GACAGGACTC GTTTCCCAGT TTGATGGCTT GATAGGCATC AGACCTCGGC CCTCCTCATT 4500
TTGGCATTTT CCGGGTTGGG GGGGCGTAGC AGTAGCACTG GAATCCTTGA TTCCTGATTT 4560
GGTTGACATT TTATTGTCTC CAGCTCGTTT CGTATTTATT GAAAATGTGT TTGCTTTGGG 4620
CTTAAAACGG TGGAACTTGT CGGAGGAATT TCTTTAACTT AACTTAAAAT TGGGTACTTA 4680
AATAATATTA AATTTAAGGT TTGATCAAAT TTTCCA 4716